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We study how birds evolved and became one of the most diverse groups of vertebrates. We study how interplay among speciation, extinction, phenotypic evolution and dispersal generated patterns of avian biodiversity over large temporal and geographic sales. We analyze molecular, phenotypic, and environmental data using phylogenetic and comparative methods to study the tempo and mode of bird evolution. Our work is highly integrative and aims for the unification of evolutionary and ecological theories of biodiversity.